It seems like Ryo is the subordinate bunny but is still getting used to that. How long have they been bonded? It sometimes takes them a little while to get the hierarchy down. Hazel, my male, has just recently bonded to Ellie, my female. Ellie is definitely top bunny but is very laid back and relaxed. Hazel was forever trying to get Ellie to groom him but while they were establishing their relationship she would consistently refuse. Sometimes she would nip him and run away when he got pushy but most of the time she would just force her head under his chin. As things progressed, he got the idea and would usually groom her. I think because Ellie is so calm and just overall a confident bunny that she warmed up to him quicker. Now she will concede to grooming him on occasion but these are always short bouts of grooming and she usually reinforces her position by having him groom her afterwards. Now Ruby and Smitty, my former rabbit pair, were different because Ruby was a typical "queen rabbit". She almost never groomed Smitty no matter how much he begged. Eventually she started rough grooming him and it turned into the very rare lick from her. Sounds like Ryo and Alex are just working things out.